Barfuss, Wolfram, Jessica Flack, Chaitanya Gokhale, Lewis Hammond, Christian Hilbe, Edward Hughes, Joel Leibo, et al. (2025) 2025. “Collective Cooperative Intelligence.”. Proceedings of the National Academy of Sciences of the United States of America 122 (25): e2319948121. doi:10.1073/pnas.2319948121.

Chen, Jiangzhuo, Baltazar Espinoza, Jingyuan Chou, Abba Gumel, Simon Levin, and Madhav Marathe. (2025) 2025. “A Simple Model of Coupled Individual Behavior and Its Impact on Epidemic Dynamics.”. Mathematical Biosciences 380: 109345. doi:10.1016/j.mbs.2024.109345.

Chen, Jiangzhuo, Baltazar Espinoza, Jingyuan Chou, Abba Gumel, Simon Levin, and Madhav Marathe. (2025) 2025. “A Simple Model of Coupled Individual Behavior and Its Impact on Epidemic Dynamics.”. Mathematical Biosciences 380: 109345. doi:10.1016/j.mbs.2024.109345.

Duporge, Isla, Zijing Wu, Zeyu Xu, Peng Gong, Daniel Rubenstein, David Macdonald, Anthony Sinclair, Simon Levin, Stephen Lee, and Tiejun Wang. (2025) 2025. “AI-Based Satellite Survey Offers Independent Assessment of Migratory Wildebeest Numbers in the Serengeti.”. PNAS Nexus 4 (9): pgaf264. doi:10.1093/pnasnexus/pgaf264.

Espinoza, Baltazar, Jiangzhuo Chen, Mark Orr, Chadi Saad-Roy, Simon Levin, and Madhav Marathe. (2025) 2025. “The Impact of Risk Compensation Adaptive Behavior on the Final Epidemic Size.”. Mathematical Biosciences 380: 109370. doi:10.1016/j.mbs.2024.109370.

Espinoza, Baltazar, Jiangzhuo Chen, Mark Orr, Chadi Saad-Roy, Simon Levin, and Madhav Marathe. (2025) 2025. “The Impact of Risk Compensation Adaptive Behavior on the Final Epidemic Size.”. Mathematical Biosciences 380: 109370. doi:10.1016/j.mbs.2024.109370.

Impalli, Isabella, Erik Bergland, Chadi Saad-Roy, Bryan Grenfell, Simon Levin, D Larsson, and Ramanan Laxminarayan. (2025) 2025. “Optimal Sampling Frequency and Site Selection for Wastewater and Environmental Surveillance of Infectious Pathogens: A Value of Information Assessment.”. PLoS Computational Biology 21 (6): e1013190. doi:10.1371/journal.pcbi.1013190.

Junquera, Victoria, José Hormaza, Daniel Rubenstein, Simon Levin, Iñaki Pérez, and Pablo Gavilán. (2025) 2025. “Severe Water Crisis in Southern Spain under Expanding Irrigated Agriculture: A Multidimensional Drought Analysis.”. Proceedings of the National Academy of Sciences of the United States of America 122 (39): e2508055122. doi:10.1073/pnas.2508055122.

Li, Yimei, Bjarke Nielsen, Simon Levin, Aartjan Velthuis, and Bryan Grenfell. (2025) 2025. “Spatio-Temporal Modelling of Influenza A Virus Infection: The Impact of Defective Interfering Particles on Type I Interferon Response.”. BioRxiv : The Preprint Server for Biology. doi:10.1101/2025.10.09.681519.

Nguyen, Maximilian, Ari Freedman, Matthew Cheung, Chadi Saad-Roy, Baltazar Espinoza, Bryan Grenfell, and Simon Levin. (2025) 2025. “The Complex Interplay Between Risk Tolerance and the Spread of Infectious Diseases.”. Journal of the Royal Society, Interface 22 (225): 20240486. doi:10.1098/rsif.2024.0486.

Qiu, Zirou, Daniel Rosenkrantz, Matthew Jackson, Simon Levin, S Ravi, Richard Stearns, and Madhav Marathe V. (2025) 2025. “Welfare Optimization for Resource Allocation With Peer Effects.”. PNAS Nexus 4 (9): pgaf298. doi:10.1093/pnasnexus/pgaf298.

Salvador, Liliana, Frederic Bartumeus, Simon Levin, and William Ryu. (2025) 2025. “Correction: ’Mechanistic Analysis of the Search Behaviour of ’ (2014), by Salvador .”. Journal of the Royal Society, Interface 22 (228): 20250557. doi:10.1098/rsif.2025.0557.

Su, Jie, Wei Wu, Denis Patterson, Simon Levin, and Jin Wang. (2025) 2025. “Revealing Physical Mechanisms of Spatial Pattern Formation and Switching in Ecosystems via Nonequilibrium Landscape and Flux.”. Advanced Science (Weinheim, Baden-Wurttemberg, Germany) 12 (26): e2501776. doi:10.1002/advs.202501776.

Crabtree, Stefani, Colin Wren, Avinash Dixit, and Simon Levin. (2024) 2024. “Influential Individuals Can Promote Prosocial Practices in Heterogeneous Societies: A Mathematical and Agent-Based Model.”. PNAS Nexus 3 (7): pgae224. doi:10.1093/pnasnexus/pgae224.

Deng, Jie, Washington Taylor, Simon Levin, and Serguei Saavedra. (2024) 2024. “On the Limits to Invasion Prediction Using Coexistence Outcomes.”. Journal of Theoretical Biology 577: 111674. doi:10.1016/j.jtbi.2023.111674.

Deng, Jie, Otto Cordero, Tadashi Fukami, Simon Levin, Robert Pringle, Ricard Solé, and Serguei Saavedra. (2024) 2024. “The Development of Ecological Systems Along Paths of Least Resistance.”. Current Biology : CB 34 (20): 4813-4823.e14. doi:10.1016/j.cub.2024.08.050.

Deng, Jie, Otto Cordero, Tadashi Fukami, Simon Levin, Robert Pringle, Ricard Solé, and Serguei Saavedra. (2024) 2024. “The Development of Ecological Systems Along Paths of Least Resistance.”. Current Biology : CB 34 (20): 4813-4823.e14. doi:10.1016/j.cub.2024.08.050.

Espinoza, Baltazar, Chadi Saad-Roy, Bryan Grenfell, Simon Levin, and Madhav Marathe. (2024) 2024. “Adaptive Human Behaviour Modulates the Impact of Immune Life History and Vaccination on Long-Term Epidemic Dynamics.”. Proceedings. Biological Sciences 291 (2033): 20241772. doi:10.1098/rspb.2024.1772.

Espinoza, Baltazar, Chadi Saad-Roy, Bryan Grenfell, Simon Levin, and Madhav Marathe. (2024) 2024. “Adaptive Human Behaviour Modulates the Impact of Immune Life History and Vaccination on Long-Term Epidemic Dynamics.”. Proceedings. Biological Sciences 291 (2033): 20241772. doi:10.1098/rspb.2024.1772.

Freedman, Ari, Justin Sheen, Stella Tsai, Jihong Yao, Edward Lifshitz, David Adinaro, Simon Levin, Bryan Grenfell, and Jessica Metcalf. (2024) 2024. “Inferring COVID-19 Testing and Vaccination Behavior from New Jersey Testing Data.”. Proceedings of the National Academy of Sciences of the United States of America 121 (17): e2314357121. doi:10.1073/pnas.2314357121.

Gibbs, Theo, Gabriel Gellner, Simon Levin, Kevin McCann, Alan Hastings, and Jonathan Levine. (2024) 2024. “When Can Higher-Order Interactions Produce Stable Coexistence?”. Ecology Letters 27 (6): e14458. doi:10.1111/ele.14458.

Grasso, Eleonora, Luke Bloy, Phillip Kaplan, Amit Bar-Or, Ann Yeh, Douglas Arnold, Sridar Narayanan, et al. (2024) 2024. “Choroid Plexus Volume in Pediatric-Onset Multiple Sclerosis.”. Neurology(R) Neuroimmunology & Neuroinflammation 11 (6): e200319. doi:10.1212/NXI.0000000000200319.

Junquera, Victoria, Maja Schlüter, Juan Rocha, Nico Wunderling, Simon Levin, Daniel Rubenstein, Jean-Christophe Castella, and Patrick Meyfroidt. (2024) 2024. “Crop Booms As Regime Shifts.”. Royal Society Open Science 11 (6): 231571. doi:10.1098/rsos.231571.

Levin, Simon, and Elke Weber. (2024) 2024. “Polarization and the Psychology of Collectives.”. Perspectives on Psychological Science : A Journal of the Association for Psychological Science 19 (2): 335-43. doi:10.1177/17456916231186614.

Levin, Simon, and Elke Weber. (2024) 2024. “Polarization and the Psychology of Collectives.”. Perspectives on Psychological Science : A Journal of the Association for Psychological Science 19 (2): 335-43. doi:10.1177/17456916231186614.

Ruttenberg, Dee, Simon Levin, Ned Wingreen, and Sarah Kocher. (2024) 2024. “Variation in Season Length and Development Time Is Sufficient to Drive the Emergence and Coexistence of Social and Solitary Behavioural Strategies.”. Proceedings. Biological Sciences 291 (2032): 20241221. doi:10.1098/rspb.2024.1221.

Ruttenberg, Dee, Simon Levin, Ned Wingreen, and Sarah Kocher. (2024) 2024. “Variation in Season Length and Development Time Is Sufficient to Drive the Emergence and Coexistence of Social and Solitary Behavioral Strategies.”. BioRxiv : The Preprint Server for Biology. doi:10.1101/2024.06.18.599518.

Ruttenberg, Dee, Simon Levin, Ned Wingreen, and Sarah Kocher. (2024) 2024. “Variation in Season Length and Development Time Is Sufficient to Drive the Emergence and Coexistence of Social and Solitary Behavioural Strategies.”. Proceedings. Biological Sciences 291 (2032): 20241221. doi:10.1098/rspb.2024.1221.

Saad-Roy, Chadi, Sinead Morris, Mike Boots, Rachel Baker, Bryan Lewis, Jeremy Farrar, Madhav Marathe V, et al. (2024) 2024. “Impact of Waning Immunity Against SARS-CoV-2 Severity Exacerbated by Vaccine Hesitancy.”. PLoS Computational Biology 20 (8): e1012211. doi:10.1371/journal.pcbi.1012211.

Segerson, Kathleen, Stephen Polasky, Marten Scheffer, Rashid Sumaila, Juan Cárdenas, Karine Nyborg, Eli Fenichel, et al. (2024) 2024. “A Cautious Approach to Subsidies for Environmental Sustainability.”. Science (New York, N.Y.) 386 (6717): 28-30. doi:10.1126/science.ado2615.

Segerson, Kathleen, Stephen Polasky, Marten Scheffer, Rashid Sumaila, Juan Cárdenas, Karine Nyborg, Eli Fenichel, et al. (2024) 2024. “A Cautious Approach to Subsidies for Environmental Sustainability.”. Science (New York, N.Y.) 386 (6717): 28-30. doi:10.1126/science.ado2615.

Cooney, Daniel, Simon Levin, Yoichiro Mori, and Joshua Plotkin. (2023) 2023. “Evolutionary Dynamics Within and Among Competing Groups.”. Proceedings of the National Academy of Sciences of the United States of America 120 (20): e2216186120. doi:10.1073/pnas.2216186120.

Dasgupta, Partha, and Simon Levin. (2023) 2023. “Economic Factors Underlying Biodiversity Loss.”. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences 378 (1881): 20220197. doi:10.1098/rstb.2022.0197.

Espinoza, Baltazar, Aniruddha Adiga, Srinivasan Venkatramanan, Andrew Warren, Jiangzhuo Chen, Bryan Lewis, Anil Vullikanti, et al. (2023) 2023. “Coupled Models of Genomic Surveillance and Evolving Pandemics With Applications for Timely Public Health Interventions.”. Proceedings of the National Academy of Sciences of the United States of America 120 (48): e2305227120. doi:10.1073/pnas.2305227120.

Nguyen, Maximilian, Ari Freedman, Sinan Ozbay, and Simon Levin. (2023) 2023. “Fundamental Bound on Epidemic Overshoot in the SIR Model.”. Journal of the Royal Society, Interface 20 (209): 20230322. doi:10.1098/rsif.2023.0322.

Nielsen, Bjarke, Chadi Saad-Roy, Yimei Li, Kim Sneppen, Lone Simonsen, Cécile Viboud, Simon Levin, and Bryan Grenfell. (2023) 2023. “Host Heterogeneity and Epistasis Explain Punctuated Evolution of SARS-CoV-2.”. PLoS Computational Biology 19 (2): e1010896. doi:10.1371/journal.pcbi.1010896.

Patterson, Denis, Simon Levin, Ann Staver, and Jonathan Touboul. (2023) 2023. “Pattern Formation in Mesic Savannas.”. Bulletin of Mathematical Biology 86 (1): 3. doi:10.1007/s11538-023-01231-7.

Saad-Roy, Chadi, Sinead Morris, Rachel Baker, Jeremy Farrar, Andrea Graham, Simon Levin, Caroline Wagner, Jessica Metcalf, and Bryan Grenfell. (2023) 2023. “Medium-Term Scenarios of COVID-19 As a Function of Immune Uncertainties and Chronic Disease.”. Journal of the Royal Society, Interface 20 (205): 20230247. doi:10.1098/rsif.2023.0247.

Saad-Roy, Chadi, Simon Levin, Bryan Grenfell, and Mike Boots. (2023) 2023. “Epidemiological Impacts of Post-Infection Mortality.”. Proceedings. Biological Sciences 290 (2002): 20230343. doi:10.1098/rspb.2023.0343.

Schrom, Edward, Ann Kinzig, Stephanie Forrest, Andrea Graham, Simon Levin, Carl Bergstrom, Carlos Castillo-Chavez, et al. (2023) 2023. “Challenges in Cybersecurity: Lessons from Biological Defense Systems.”. Mathematical Biosciences 362: 109024. doi:10.1016/j.mbs.2023.109024.

Sood, Mansi, Anirudh Sridhar, Rashad Eletreby, Chai Wu, Simon Levin, Osman Yağan, and Vincent Poor. (2023) 2023. “Spreading Processes With Mutations over Multilayer Networks.”. Proceedings of the National Academy of Sciences of the United States of America 120 (24): e2302245120. doi:10.1073/pnas.2302245120.

Tian, Yurun, Anirudh Sridhar, Chai Wu, Simon Levin, Kathleen Carley, Vincent Poor, and Osman Yağan. (2023) 2023. “Role of Masks in Mitigating Viral Spread on Networks.”. Physical Review. E 108 (1-1): 014306. doi:10.1103/PhysRevE.108.014306.

Traulsen, Arne, Simon Levin, and Chadi Saad-Roy. (2023) 2023. “Individual Costs and Societal Benefits of Interventions During the COVID-19 Pandemic.”. Proceedings of the National Academy of Sciences of the United States of America 120 (24): e2303546120. doi:10.1073/pnas.2303546120.

Xu, Li, Denis Patterson, Simon Levin, and Jin Wang. (2023) 2023. “Non-Equilibrium Early-Warning Signals for Critical Transitions in Ecological Systems.”. Proceedings of the National Academy of Sciences of the United States of America 120 (5): e2218663120. doi:10.1073/pnas.2218663120.

Xu, Li, Denis Patterson, Simon Levin, and Jin Wang. (2023) 2023. “Non-Equilibrium Early-Warning Signals for Critical Transitions in Ecological Systems.”. Proceedings of the National Academy of Sciences of the United States of America 120 (5): e2218663120. doi:10.1073/pnas.2218663120.

Chapin, Stuart, Elke Weber, Elena Bennett, Reinette Biggs, Jeroen van den Bergh, Neil Adger, Anne-Sophie Crépin, et al. (2022) 2022. “Earth Stewardship: Shaping a Sustainable Future through Interacting Policy and Norm Shifts.”. Ambio 51 (9): 1907-20. doi:10.1007/s13280-022-01721-3.

Cooney, Daniel, Dylan Morris, Simon Levin, Daniel Rubenstein, and Pawel Romanczuk. (2022) 2022. “Social Dilemmas of Sociality Due to Beneficial and Costly Contagion.”. PLoS Computational Biology 18 (11): e1010670. doi:10.1371/journal.pcbi.1010670.

Cooney, Daniel, Fernando Rossine, Dylan Morris, and Simon Levin. (2022) 2022. “A PDE Model for Protocell Evolution and the Origin of Chromosomes via Multilevel Selection.”. Bulletin of Mathematical Biology 84 (10): 109. doi:10.1007/s11538-022-01062-y.

Fahimipour, Ashkaan, Fanqi Zeng, Martin Homer, Arne Traulsen, Simon Levin, and Thilo Gross. (2022) 2022. “Sharp Thresholds Limit the Benefit of Defector Avoidance in Cooperation on Networks.”. Proceedings of the National Academy of Sciences of the United States of America 119 (33): e2120120119. doi:10.1073/pnas.2120120119.

Fischer, Ilan, Daniel Rubenstein, and Simon Levin. (2022) 2022. “Vaccination-Hesitancy and Global Warming: Distinct Social Challenges With Similar Behavioural Solutions.”. Royal Society Open Science 9 (6): 211515. doi:10.1098/rsos.211515.